Selected publications

For a complete list of publications please visit Google Scholar

Image by CHUTTERSNAP
Late Cretaceous origins for major nightshade lineages from total-evidence timetree analysis

Late Cretaceous origins for major nightshade lineages from total-evidence timetree analysis

Gonzalez-Ramirez, Ixchel S.; Deanna, Rocio; Smith, Stacey D.

Annals of Botany 2026

We implement a bayesian total evidence dating analysis to infer the divergence times of Solanaceae. The use of this integrative model allowed us to simultaneously explore fossil position and infer divergence times while accounting for topological uncertainty.

nightshades timetree total evidence dating
Spatial phylogenetics with continuous data: an application to California bryophytes

Spatial phylogenetics with continuous data: an application to California bryophytes

Kling, Matthew M.; Gonzalez-Ramirez, Ixchel S.; Carter, Benjamin; Borokini, Israel; Mishler, Brent D.

Systematic Botany 2025

In this paper we present smoothed spatial phylogenetics metrics, that directly use taxa probability of occurrence without the need of occurrence thresholds. We apply these metrics to a dataset of California bryophytes to uncover diversity patterns across this biodiversity hotspot.

spatial phylogenetics bryophytes California
Typification of *Fimbraria californica* (*Calasterella californica*, Aytoniaceae)

Typification of Fimbraria californica (Calasterella californica, Aytoniaceae)

Gonzalez-Ramirez, Ixchel S.; Long, David G.; Briscoe, Laura; Mishler, Brent D.

Madroño 2025

We clarify the type specimen of Calasterella californica. This involved clarifying a previous lectotypification by Evans, and designing an epitype from a recently collected specimen with genetic information available.

taxonomy typification Calasterella
A genome assembly for the California endemic liverwort *Calasterella californica*

A genome assembly for the California endemic liverwort Calasterella californica

Gonzalez-Ramirez, Ixchel S.; Escalona, Merly; Miller, Courtney; Chumchim, Noravit; Marimuthu, Mohan P.A.; Nguyen, Oanh; Song, Michael J.; Mishler, Brent D.

Journal of Heredity 2025

Funded by the California Conservation Genomics Project, we assembled de novo the genome of Calasterella californica, a West Coast endemic and xerophytic liverwort.

genome assembly liverworts Calasterella
Unveiling nature's sunscreen: the potential role of liverwort scales in UV protection

Unveiling nature’s sunscreen: the potential role of liverwort scales in UV protection

Jain, Riya; Shahryari, Atra; Gonzalez-Ramirez, Ixchel S.

Madroño 2024

Various xerophytic liverworts role into their ventral scales when dessecated. Often, these liverworts also have purple scales. In this paper, we test whether these red ventral scales differentially absorbe UV-radiation. This constitutes the first step towards understanding the potential role of ventral scales on adaptation to xeric environments.

liverworts UV protection scales